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SYSTEMATIC PALEONTOLOGY

DINOSAURIA Owen, 1841

THEROPODA Marsh, 1881

TETANURAE Gauthier, 1986

TYRANNOSAURIDAE Osborn, 1906
Tyrannosaurus rex Osborn, 1905

MATERIAL: The specimen was collected in Carter County, Montana in the upper middle portion of the Hell Creek Formation (Maastrichtian).

DESCRIPTION: The bones have well-preserved periosteal surfaces with osteological details evident on many of the bones. There is some crushing and breakage of some elements. Parts of the skull and skeleton were recovered. Cranial material comprises the quadrate, postorbital, nasal, and the posterior mandibular elements the right and left articular, a prearticular, and an angular. The hindlimb elements recovered were portions of the femur, a nearly complete tibia, a complete metatarsal IV and a right pes distal phalanx. Cervical and caudal ribs, several caudal vertebrae and two chevrons were also recovered. Partial vertebrae were found but not identified as to level in the vertebral column. “Titus” is the 16th most complete of all Tyrannosaurus rex specimens granted museum accession numbers Elements are in the size range of other, more complete, adult T. rex specimens.

DIAGNOSIS: The specimen is referred to Tyrannosaurus rex based on the following characters: Fused nasals and Metatarsal IV (plus see new characters described by Loewen et al., 2013). This referral is also confirmed by size comparison to adult T. rex.

NICKNAME: “Titus” from Shakespeare’s play: TITUS ANDRONICUS— “Blood and revenge are hammering in my head”— ACT 2 SCENE 3.

DISCOVER THE FULL TITUS REPORT

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